I've begun thinking about the various aliens in the universe of our beloved MMO. And I mean -space aliens-, not beings from other dimensions. So far, I've compiled this listing, with what I know and some of what I surmise....
Kheldians - Formless energy beings from another galaxy, their natural forms look like floating puffballs of energy, blue-white in color. They can manipulate energy in various ways, thus leading to their innate ability to shapeshift and merge with organic hosts. They normally have control of light and energy, and are generally noble, peaceful creatures. Khledians without a host have a lifespan of about 10 Earth years, but by moving from host to host, they become functionally immortal (the Warshade Shadowstar is about 3000-5000 years old, if rumors are true).
Nictus are evil Khledians who have used advanced science to transform themselves and their powers. Black-purple in color, they now manipulate gravity and dark matter in a fashion similar to standard Kheldians. The lifespan of an unjoined Nictus (and Warshades) is about 5 years. Joined Nictus are, like Kheldians, unaging while in a host.
The Nictus once ruled their race with an iron fist (metaphorically-speaking), until the normal Kheldians rose up and cast them down. The Nictus fled, and those that hunted them became known as the Peacebringers. What resulted was a running war across the universe, with the Nictus forcefully possessing various alien races to hide in, and the Peacebringers finding them and driving them out. The war has now come to Earth, with the Nictus having been here for centuries, if not millennia, and the Peacebringers just recently arriving. The Nictus have joined forces with the Council as part of their plans to conquer Earth.
The Warshades are Nictus that have reformed and become heroes, enjoying a generally peaceful, if uncomfortable, alliance with the Peacebringers to battle the Nictus.
Kurukt/Ruktur - Large, heavily armored creatures, sometimes described as "lobsters" by humans. The Kurukt evolved on the surface of a neutron star, thus making them incredibly strong and nearly indestructable. The Ruktur, likewise, come from the surface of a pulsar, gaining similar physical might. The Peacebringers and Nictus once used these creatures as hosts and allies, and their physical patterns are retained by them today as the "White Dwarf" and "Black Dwarf" forms, respectively.
Despite evolving on the surfaces of two different stars, it's clear the Kurukt and Ruktur were at some point one species. After the Peacebringer-Nictus war moved on, it is not known what happened to these races.
Mefnanim/Hulnanim - These two alien races are, likewise, clearly branches of the same race. Squid-like in appearance, and naturally bouyant, these creatures evolved in the atmophere of a gas giant. The Mefnanim lived in the energy-charged upper atmosphere, while the Hulnanim lived in the dark depths of the lower atmosphere. Both species had the ability to project blasts of energy or negative energy. As with the "lobsters," the "squids" became allies and pawns of the Peacebringers and Nictus, and their bodies and powers are remembered by the Kheldians today as the "Bright Nova" and "Dark Nova" forms. When the Kheldian war left their world, it is not revealed what became of the "squids."
Maaxigada'arzildilo - The only known representative of this alien race is the hero Dillo. Little is known about them, beyond that their world was assaulted by the Shivans, and the "Maaxi" were forced into exile. Dillo came to Earth to warn humanity of the Shivan threat only to find he was too late. They were already here.
Dillo wears some sort of containment suit, that, if breached, leaves him severely weakened and unable to fight until it is repaired and re-pressurized. This suggests that the Maaxi cannot survive in Earth's atmosphere unaided, possibily to not being able to breath oxygen, or that Earth's air pressure is unsuitable for Maaxi life. They are humanoid, and somewhat squat, judging from Dillo's appearance, but beyond that we cannot be certain what they look like. They appear to be a friendly species.
Shivans - The scouts of the "Coming Storm," these creatures seem to be living goo. They ride inside meteors launched by the Coming Storm, and are perfectly capable of surviving atmospheric entry and landing. This ooze combines with whatever unliving matter is nearby to form skeletal structures for themselves, so that they can move in the planet's gravity and atmosphere. On Bloody Bay, the Shivans combined with the buried corpses of the island's numerous old cemetaries, using skeletons to provide physical support for themselves. When they struck Galaxy City, the Shivans used the rubble created by their meteors' impacts to form bodies of concrete and steel rebar. Both types seem to explode like water balloons when they die.
It is unclear if Shivans are intelligent or sentient. They cannot speak, lacking mouths of any sort, and it is unknown if telepathic communication has ever succeeded. They are hostile and violent, and can project blasts of radiation at their enemies. It is possible that the Coming Storm simply unleashes them as "beasts of war" to soften up potential targets.
"The Coming Storm" - Almost nothing is yet known of them, other than they are hostile, use the Shivans as some sort of slave soldiers or hired guns, and are aiming at Primal Earth next. They devastated the homeworld of the Maaxigada'arzildilo, driving the survivors into exile.
Conjecture: If the Kheldian known as Twilight's Son is to be believed, in the far future the Coming Storm wiped out his entire race (using them as power sources for their ships), with himself being the last one. I don't know if this was in fact the Coming Storm that did this (the descriptions never name TS's "masters"), but seems reasonable, as Twilight's Son is an Ouroboros Mender, and the Menders show considerable wariness about the Coming Storm.