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If I ran the zoo, here's what I'd do...
The path to "Incarnate" status should be a Hero's Quest. Not unlike Heracles, there should be a number of tasks to accomplish, all of them difficult, some which require a team, some which must be done solo. Perhaps even a task which you must accomplish as an exemplar. All of the tasks in the arc should have one thing in common: the quester must be directly involved. This can't be done for you. (Though I'm not opposed to the idea of one task that must be done for you.)
Each task should test the quester in a different area: strength, courage, humility, self-sacrifice, etc. All the virtues an Incarnate should possess.
Failure of one or two tasks would not lead to failure of the quest, per se, but may shape the final outcome: the lacking virtues could constitute a weakness of the Incarnate. There should be a critical mass, however, at which point the quest is failed and cannot be repeated until you've earned the right. Yes, it's harsh, but Incarnate must be earned. There should be no easy way to it.
I would like paths to the end to vary. Unlike current story arcs in which everyone runs the same missions in the same order, let there be 3-4 tasks which can be accomplished for each virtue. This would lead to high variation the quest from player to player.
A Hero's Quest always involves some manner of sacrifice. This should be no different. The quester must be required to relinquish something of great value. Not influence, but something that would really count. An important power, perhaps, or several levels.
Completion of the quest should incur a reward consumate with the risk taken, and would elevate the character into a new realm of possibilities and dangers heretofore unimagined. It's a whole new ballgame.
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Excellent idea, but Statesman (the dev) has said to ideas like this that having quests and what not that invoke thought and challege would be easily found out and the subsequently exploited. I like the idea, but it would just take one lazy player to see the most powerful incarnate and then go to the boards and see what missions/quests were done and how to do them in order to get the most powerful resultant character.
If only there was an expansion that implemented some sort of delightful algorithm for players that like to work at the game and have fun with it rather than people who obviously don't.